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Dans les tunnels secrets du métro, Isis découvre le clan des parias, des souris exclues du Louvre. Avec l'aide d'Antoinette, la doyenne des souris qui connaît tous les mystères du musée, elle va porter secours à Ésope qui a été capturé. Pendant ce temps, Milo, Victoire et Bastien s'introduisent dans le Louvre pour aider les souris à mettre un terme à la guerre lancée par Xanthos...
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